List of Subjects



Crop Ecology

lessons in hour (lecture + practice + home) / week =2+2+4

Credit: 6

PhD course

Zoltán Nagy


The course focuses on production processes in terrestrial agro-ecosystems. Topics include discussion of production limiting environmental factors, related transport processes (air and soil), investigation of production processes from leaf to ecosystem scale utilising concepts from the fields of terrestrial plant ecology, plant ecophysiology and micrometeorology. Emphasis will be given to aspects of water limited production considering concepts and methods employed in the field of plant water relations from leaf to field scale. Methods applied to follow/characterize phenological development and plant stress states will be introduced and demonstrated in lab and field practicals.


The course will use chapters from the textbooks in the below list:


David J. Connor, Robert S. Loomis (2011): Crop Ecology Productivity and Management in Agricultural Systems Cambridge University Press, 538 pp.

Chapin FS, Matson PA, Mooney HA (2002): Principles of terrestrial ecosystem ecology. Springer. 328 pp.

Monteith J., Unsworth M. (2007):  Principles of Environmental Physics, Third Edition, Elsevier, 418 pp.


An introduction to data analysis using R

lessons in hour (lecture + practice + home) / week =2+2+4

Credit: 6

PhD course

János Balogh

This course provides an introduction to statistics ideal for PhD students in ecology or related fields. The course will consist of computer practicals using the free software package R for statistics and graphics.

Topics covered include proper organisation of the workspace, reading and writing data files, using R as a calculator, using logic operators, manipulating data frames, summarising and aggregating data, programming ‘ifelse’ statements, loops, short routines, handling time fields in data frames, drawing and customising graphs.

Starting work on own data sets is the major goal of the course.


Content based English in Plant Science

PhD course

lessons in hour (lecture + practice + home) / week =2+2+8

Credit: 8

Sándor Fekete, János Balogh


The course is designed to give PhD students the opportunity to develop their technical language skills in a range of areas such as giving presentations, taking part in scientific formal and informal discussions, reading technical texts and writing scientific articles.

These aims and objectives are achieved by studying some of the relevant chapters of modern plant physiology and molecular biology such as photosynthesis, xylem and phloem transport, plant growth regulators/hormones, gene expression regulation in response to stress as well as some key concepts in ecology like biogeochemical cycles. The chapters are dealt with in English throughout the course with special emphasis on key vocabulary and language patterns in both formal and informal technical language and the students are encouraged to take an active part in both the presentation of new contents and the discussions following them.



  1. General overview of the cell, cellular organelles and their functions
  2. Cellular respiration: How the cell makes ATP
  3. Turning light energy into chemical energy, electron transport chain and Calvin cycle.
  4. Water conduction in plants
  5. Phloem transport
  6. Carbon cycle, nitrogen cycle
  7. Griffith, Avery, Hershey &Chase experiments, DNA replication
  8. Eukaryotic mRNA synthesis/transcription and protein synthesis/translation
  9. Prokaryotic gene expression regulation — lactose operon, eukaryotic gene expression regulation
  10. Plant hormones: auxine molecular mechanism of action, gibberellins, cytokinins, ABA, ethylene
  11. Plant stress, regulation of stress gene expression in plants
  12. The conquest of land / Moss life cycle / Fern life cycle / Gymnosperm life cycel/ Angiosperm life cycle and double fertilisation